\[\begin{array}{rcl} {d A \over d t} & = & k_{21} A \: + \: k_{12} L V \\ {d L \over d t} & = & {K_{21} A \over V} \: - \: \left ( k_{12} \: + \: k_{el} \right ) L \: - \: k_{on} L R \: + \: k_{off} P \\ {d R \over d t} & = & k_{syn} \: - \: k_{deg} R \: - \: k_{on} L R \: + \: k_{off} P \\ {d P \over d t} & = & k_{on} L R \: - \left ( k_{off} + k_{int} \right ) P \end{array}\]